Well put it this way.....you have 3 large ranges of carb calibration....Idle, WOT, and everything in between......If one of those ranges isn't right, then it'll cause hesitation and bog....If the pilots or idle range is running rich or lean, then when you go into another range, it takes a second for the air/ fuel to change over to a different mixture that the motor likes better.....thats where your hesitation comes from....If the pilots are perfect and mains are perfect, and the needles aren't, then again, you'll have a hesitation when coming from 1/4-3/4 to wot.....so you can see that not only mains are important, but everything works together...All 3 ranges must be perfect to get great response...This takes hours, but well worth the hours of great riding after.....
How often are you totally off or totally full throttle??? 20%?? The other 80% is in the needle range, so now you see why the needles make such a huge difference...Next time you ride, think about how far the throttle is being pushed, you'll be amazed how you hardly ever open it up fully.....